Recent Publications

In press

  • Merlo, E.M., Milligan, K.A., Sheets, N.B., Neufeld, C.J., Eastham, T.M., Estores-Pacheco, K.A.L., Steinke, D., Hebert, P,D.N., Valdés, A., & Wyeth, R.C. In press. Range extension for the region of sympatry between the nudibranchs Hermissenda opalescens and Hermissenda crassicornis in the Northeastern Pacific. FACETS.
  • Medrano, S., Krug, P.J., Gosliner, T.M., Kumar, A.B., & Valdés, A. In press. Systematics of Polybranchia Pease, 1860 (Mollusca: Gastropoda: Sacoglossa) based on molecular and morphological data. Zoological Journal of the Linnean Society.
  • Krug, P.J., Berriman, J.S., & Valdés, A. In press. Phylogenetic systematics of the shelled sea slug genus Oxynoe Rafinesque, 1814 (Heterobranchia: Sacoglossa), with integrative descriptions of seven new species. Invertebrate Systematics.


  • Berriman, J.S., Ellingson, R.A., Awbrey, J.D., Rico, D.M., Valdés, A., Wilson, N.G., Aguilar, A., Herbert, D.G., Hirano, Y.M., Trowbridge, C.D., Krug, P.J. 2018. A biting commentary: Integrating tooth characters with molecular data doubles known species diversity in a lineage of sea slugs that consume “killer algae.” Molecular Phylogenetics and Evolution 126: 356-370 doi:


  • Valdés, A., Breslau, E., Padula, V., Schrödl, M., Camacho, Y., Malaquias, M., Alexander, J., Bottomley, M., Vital, X., Hooker, Y. & Gosliner, T.M. 2017. Molecular and morphological systematics of Dolabrifera Gray, 1847 (Mollusca, Gastropoda, Heterobranchia, Aplysiomorpha). Zoological Journal of the Linnean Society doi:
  • McCarthy, J.B., Krug, P.K., & Valdés, A. 2017. Integrative systematics of Placida cremoniana (Trinchese, 1892) (Gastropoda, Heterobrachia, Sacoglossa) reveals multiple pseudocryptic species. Marine Biodiversitydoi:10.1007/s12526-017-0812-2
  • Goodheart, J., Bazinet, A.L., Valdés, A., Collins, A.G., & Cummings, M. 2017. Prey preference follows phylogeny: Evolutionary dietary patterns within the marine gastropod group Cladobranchia (Gastropoda: Heterobranchia: Nudibranchia). BMC Evolutionary Biology 17:221 doi:10.1186/s12862-017-1066-0
  • Hoover, C.A., Padula, V., Schrödl, M., Hooker, Y. & Valdés, A. 2017. Integrative taxonomy of the Felimare californiensis and Felimare ghiselini species complex (Mollusca, Gastropoda, Nudibranchia) and description of a new species from Peru. Journal of Molluscan Studies 83: 461-475 doi:
  • Valdés, A., Gosliner, T.M. & Warén, A. 2017. A new species of Parvaplustrum Powell, 1951 (Gastropoda, Heterobranchia: Aplustridae) from the northeastern Pacific. The Nautilus 131: 97–100.
  • Uribe, R., Sepúlveda, F., Goddard, J.H.R. & Valdés, A. 2017. Integrative systematics of the genus Limacia O. F. Müller, 1781 (Mollusca, Gastropoda, Nudibranchia, Polyceridae) in the Eastern Pacific. Marine Biodiversity doi:10.1007/s12526-017-0676-5
  • Garabedian, K., Malaquias, M.A.E., Crocetta, F., Zenetos, A., Kavadas, S., & Valdés, A. 2017. Haminoea orteai Talavera, Murillo and Templado, 1987 (Mollusca: Gastropoda: Heterobranchia: Cephalaspidea), a widespread species in the Mediterranean and northeastern Atlantic. Cahiers de Biologie Marine 58: 107-113 doi:10.21411/CBM.A.19015AD4
  • Breslau, E., Valdés, A. & Chichvarkhin, A. 2017. A new cryptic species of Melanochlamys (Gastropoda: Heterobranchia: Cephalaspidea) from the Northwestern Pacific. American Malacological Bulletin 34: 103-108 doi: 10.4003/006.034.0206


  • Ekimova, I., Valdés, A., Schepetov, D., Chichvarkhin, A. 2016. Was Gordon Robilliard right? Integrative systematics suggests that Dendronotus diversicolor Robilliard, 1970 is a valid species. Canadian Journal of Zoology 94: 793-799 doi:10.1139/cjz-2016-0096
  • Lindsay, T., Kelly, J., Chichvarkhin, A., Craig, S., Kajihara, H., Mackie, J & Valdés, A. 2016. Changing spots: pseudocryptic speciation in the North Pacific dorid nudibranch Diaulula sandiegensis (Cooper, 1862) (Gastropoda: Heterobranchia). Journal of Molluscan Studies 82: 564-574 doi:10.1093/mollus/eyw026
  • Chichvarkhin, A.Y., Ekimova, I.A., Chalenko, K.P., Schepetov, D., Chichvarkhina, O.V. & Valdés. A. 2016. Placida babai (Mollusca: Gastropoda: Sacoglossa) from Russian waters of the Sea of Japan. Bulletin of the Russian Far East Malacological Society 20: 44–56.
  • Moustafa, A.Y., Golestani, H. & Valdés, A. 2016. A review of the genus Berthellina (Mollusca: Heterobranchia: Pleurobranchidae) in the Red Sea based on molecular and anatomical data. Marine Biodiversity doi:10.1007/s12526-016-0558-2
  • Krug, P.J., Vendetti, J.E. & Valdés, A. 2016. Molecular and morphological systematics of Elysia Risso (Heterobranchia: Sacoglossa) from the Caribbean region. Zootaxa 4148: 1-137 doi:10.11646/zootaxa.4148.1.1
  • Valdés, A., Cadien, D.B. & Gosliner, T.M. 2016. Philinidae, Laonidae and Philinorbidae (Cephaslaspidea: Philinoidea) from the Northeastern Pacific Ocean and the Beaufort Sea (Arctic Ocean). Zootaxa 4147: 501-537 doi:10.11646/zootaxa.4147.5.1
  • Goodheart, J.A., Ellingson, R.A., Vital, X.G., Galvão Filho, H., McCarthy, J.B., Medrano, S.M., Bhave, V.J., García-Méndez, K., Jiménez, L.M., López, G., Hoover, C.A., Awbrey, J.D., DeJesus, J.M., Gowacki, W., Krug, P.J. & Valdés, A. 2016. Identification guide to the heterobranch sea slugs (Mollusca: Gastropoda) from Bocas del Toro, Panama. Marine Biodiversity Records 9: 56 doi:10.1186/s41200-016-0048-z
  • Valdés, A., Medrano, S.M. & Bhave, V.J. 2016. A new species of Cuthona (Heterobranchia: Nudibranchia: Tergipedidae) from the Caribbean Sea. The Nautilus 130: 72–78.
  • Araya, J.F. & Valdés, A. 2016. Shallow water heterobranch sea slugs (Gastropoda: Heterobranchia) from the Región de Atacama, northern Chile. PeerJ 4:e1963
  • Lindsay, T. & Valdés, A. 2016. The model organism Hermissenda crassicornis (Gastropoda: Heterobranchia) is a species complex. PLoS ONE 11: e0154265 doi:10.1371/journal.pone.0154265
  • Valdés, A., Murillo, F.J., McCarthy, J.B. & Yedinak, N. 2016. New deep-water records and species of North Atlantic nudibranchs (Mollusca, Gastropoda: Heterobranchia) with the description of a new species. Journal of the Marine Biological Association of the United Kingdom 97: 303-319 doi:


  • Valdés. A. & McLean, J.H. 2015. On two abyssal species of Scaphandridae G. O. Sars, 1878 (Gastropoda, Cephalaspidea) from the eastern Pacific. The Nautilus 129: 118-125.
  • Gosliner, T.M., Valdés, A. & Behrens, D.W. 2015. Nudibranch and Sea Slug Identification - Indo-Pacific. New World Publications, Jacksonville, Florida.
  • Krug, P.J., Vendetti, J.E., Ellingson, R.A., Trowbridge, C.D., Hirano, Y.M., Trathen, D.Y., Rodriguez, A.K., Swennen, C., Wilson, N.G., & Valdés, A. 2015. Species selection favors dispersive life histories in sea slugs but higher per-offspring investment drives shifts to short-lived larvae. Systematic Biology 64: 983-999 doi:10.1093/sysbio/syv046
  • Hulett, R.E., Mahguib, J., Gosliner, T.M. & Valdés, A. 2015. Molecular evaluation of the phylogenetic position of the enigmatic species Trivettea papalotla (Bertsch et al. 2009) (Mollusca, Nudibranchia). Invertebrate Systematics 29: 215-222 doi:10.1071/IS15002
  • Hoover, C. Lindsay, T., Goddard, J.H.R., & Valdés, A. 2015. Seeing double: Pseudocryptic diversity in the Doriopsilla albopunctata-Doriopsilla gemela species complex of the north-eastern Pacific. Zoologica Scripta 44: 612-631 doi:10.1111/zsc.12123
  • Goddard, J.H.R. & Valdés, A. 2015. Reviving a cold case: two northeastern Pacific dendrodorid nudibranchs reassessed (Gastropoda: Opisthobranchia). The Nautilus 129: 31-42.
  • Goodheart, J., Camacho-García, Y., Padula, V., Schrödl, M., Cervera, J.L., Gosliner, T.M. & Valdés, A. 2015. Systematics and biogeography of Pleurobranchus Cuvier, 1804, sea slugs (Heterobranchia: Nudipleura: Pleurobranchidae). Zoological Journal of the Linnean Society 174: 322-362 doi:10.1111/zoj.12237
  • Mahguib, J. & Valdés, A. 2015. Molecular investigation of the phylogenetic position of the polar nudibranch Doridoxa (Mollusca, Gastropoda, Heterobranchia). Polar Biology 38: 1369-1377 doi:10.1007/s00300-015-1700-5


  • Behrens, D.W., Ornelas, E. & Valdés, A. 2014. Two new species of Velutinidae Gray, 1840, from the Northern Pacific with a preliminary molecular phylogeny of the family. The Nautilus 128: 114-121.
  • Espinoza, E., DuPont, A. & Valdés, A. 2014. Molecular data reveal an undescribed cryptic species of Costasiella Pruvot-Fol, 1951 (Euthyneura: Sacoglossa) in the Bahamas. American Malacological Bulletin 32: 173-182 doi:10.4003/006.032.0208
  • De Oliveira Saad, L., Magenta Cunha, C., Delevati Colpo. K., Valdés, A. 2014. It’s not what it looks like: Molecular data fails to substantiate morphological differences in two sea hares (Mollusca, Heterobranchia, Aplysiidae) from southern Brazil. Helgoland Marine Research 68: 523-530 doi:10.1007/s10152-014-0408-1
  • Cooke, S., Hanson, D., Hirano, Y., Ornelas-Gatdula, E., Gosliner, T. M., Chernyshev, A. & Valdés, A. 2014. Cryptic diversity of Melanochlamys sea slugs (Gastropoda, Aglajidae) in the North Pacific. Zoologica Scripta 43: 351-369 doi:10.1111/zsc.12063
  • Churchill, C., Valdés, A., & Ó Foighil, D. 2014. Molecular and morphological systematics of neustonic nudibranchs (Mollusca, Gastropoda, Glaucidae, Glaucus) with descriptions of three new cryptic species. Invertebrate Systematics 28: 174–195 doi:10.1071/IS13038
  • Churchill, C., Valdés, A., & Ó Foighil, D. 2014. Afro-Eurasia and the Americas present barriers to gene flow for the cosmopolitan neustonic nudibranch Glaucus atlanticus. Marine Biology 161: 899–910 doi:10.1007/s00227-014-2389-7
  • Carmona, L., Lei, B.R., Pola, M., Gosliner, T.M., Valdés, A. & Cervera, J.L. 2014. Untangling the Spurilla neapolitana (Delle Chiaje, 1841) species complex: A review of the genus Spurilla Bergh, 1864 (Mollusca: Nudibranchia: Aeolidiidae). Zoological Journal of the Linnean Society 170: 132–15 doi:10.1111/zoj.12098
  • Camacho-García, Y.E, Ornelas-Gatdula, E. Gosliner, T.M. & Valdés, A. 2014. Phylogeny of the family Aglajidae (Pilsbry, 1895) (Opisthobranchia: Cephalaspidea) inferred from mtDNA and nDNA. Molecular Phylogenetics and Evolution 71: 113-126 doi:10.1016/j.ympev.2013.11.010
  • Santander, M. & Valdés, A. 2014. Genetic structure of Polycera alabe and P. atra (Mollusca: Opistobranchia: Nudibranchia) in the Pacific coast of North America. Bulletin of the Southern California Academy of Sciences 112: 176-184


  • Valdés, A., Alexander, J., Crocetta, F., Yokeş, M.B., Giacobbe, S., Poursanidis, D., Zenetos, A., Cervera, J.L., Caballer, M., Galil, B.S., Schembri, P.J. 2013. The origin and dispersal pathway of the spotted sea hare Aplysia dactylomela (Mollusca: Opisthobranchia) in the Mediterranean Sea. Aquatic Invasions 8: 427–436 doi:10.3391/ai.2013.8.4.06
  • Hanson, D., Hirano, Y. & Valdés, A. 2013. Population genetics of Haminoea (Haloa) japonica Pilsbry, 1895, a widespread non-native sea slug (Mollusca: Opisthobranchia) in North America and Europe. Biological Invasions 15: 395-406. doi:10.1007/s10530-012-0294-x
  • Goddard, J.H.R., Schaefer, M.C., Hoover, C. & Valdés, A. 2013. The regional extinction of a conspicuous dorid nudibranch (Mollusca: Gastropoda) in California. Marine Biology 160: 1497-1510. doi:10.1007/s00227-013-2204-x
  • Crocetta, F., Macali, A., Furfaro, G., Cooke, S., Villani, G. & Valdés, A. 2013. Alien molluscan species established along the Italian shores: an update, with discussions on some Mediterranean “alien species” categories. ZooKeys 277: 91-108. doi:10.3897/zookeys.277.4362
  • Alexander, J. & Valdés, A. 2013. The ring doesn’t mean a thing: Molecular data suggests a new taxonomy for two Pacific species of sea hares (Mollusca, Opisthobranchia, Aplysiidae). Pacific Science 67: 283-294. doi:10.2984/67.2.10
  • Goodheart, J. & Valdés, A. 2013. Re-evaluation of the Doriopsilla areolata Bergh, 1880 (Mollusca: Opisthobranchia) subspecies complex in the eastern Atlantic Ocean and its relationship to South African Doriopsilla miniata (Alder & Hancock, 1864) based on molecular data. Marine Biodiversity 43: 113-120. doi:10.1007/s12526-012-0136-1
  • Gonzalez, L., Hanson, D. & Valdés, A. 2013. Molecular divergence between two sympatric species of Dondice (Mollusca: Nudibranchia) with distinct feeding specializations. Journal of the Marine Biological Association of the United Kingdom 93: 1887-1893. doi:10.1017/S0025315413000465
  • Valdés, A., Ornelas-Gatdula, E. & DuPont, A. 2013. Color pattern variation in a shallow water species of opisthobranch mollusk. Biological Bulletin 224: 35-46.
  • Churchill, C., Alejandrino, A., Valdés, A., & Ó Foighil, D. 2013. Parallel changes in genital morphology delineate cryptic diversification in planktonic nudibranchs. Proceedings of the Royal Society Biological Sciences 280: 1765 20131224. doi:10.1098/rspb.2013.1224
  • Espinoza, E., DuPont, A. & Valdés, A. 2013. A tropical Atlantic species of Melibe Rang, 1829 (Mollusca, Nudibranchia, Tethyiidae). ZooKeys 316: 55-66. doi:10.3897/zookeys.316.5452
  • Valdés, A., Blanchard, L. & Marti, W. 2013. Caught naked: First report a nudibranch sea slug attacked by a cone snail. American Malacological Bulletin 31: 337-338. doi:
  • Hanson, D., Cooke, S., Hirano, Y., Malaquias, M.A.E., Crocetta, F. & Valdés, A. 2013. Slipping through the cracks: The taxonomic impediment conceals the origin and dispersal of Haminoea japonica, an invasive species with impacts to human health. PLoS ONE 8(10): e77457. doi:10.1371/journal.pone.0077457

Full list


Lab Location: Building 4 Room: 2-405